Earthworms are a good example of soil-dwelling detritivores

Detritivores, also known as detritophages or detritus feeders or detritus eaters or saprophages, are heterotrophs that obtain nutrients by consuming detritus (decomposing plant and animal parts as well as organic fecal matter).^[1] By doing so, they contribute to decomposition and the nutrient cycles. They should be distinguished from other decomposers, such as many species of bacteria, fungi and protists, which are unable to ingest discrete lumps of matter, but instead live by absorbing and metabolizing on a molecular scale. However, the terms detritivore and decomposer are often used interchangeably.

Fungi are the primary decomposers in most environments, illustrated here Mycena interrupta. Only fungi produce the enzymes necessary to decompose lignin, a chemically complex substance found in wood.

Detritivores are an important aspect of many ecosystems. They can live on any soil with an organic component, including marine ecosystems, where they are termed interchangeably with bottom feeders.

Typical detritivorous animals include millipedes, woodlice, dung flies, slugs, many terrestrial worms, sea stars, sea cucumbers, fiddler crabs, and some sedentary polychaetes such as amphitrites (Amphitritinae, worms of the family terebellidae) and other terebellids.

Scavengers are typically not thought to be detritivores, as they generally eat large quantities of organic matter, but both detritivores and scavengers are specific cases of consumer-resource systems.^[2] The eating of wood, whether live or dead, is known as xylophagy. Τhe activity of animals feeding only on dead wood is called sapro-xylophagy and those animals, sapro-xylophagous.

Ecology [edit]

In food webs, detritivores generally play the role of decomposers. Detritivores are often eaten by consumers and therefore commonly play important roles as recyclers in ecosystem energy flow and biogeochemical cycles.

Many detritivores live in mature woodland, though the term can be applied to certain bottom-feeders in wet environments. These organisms play a crucial role in benthic ecosystems, forming essential food chains and participating in the nitrogen cycle.^[3]

Fungi, acting as decomposers, are important in today's terrestrial environment. During the Carboniferous period, fungi and bacteria had yet to evolve the capacity to digest lignin, and so large deposits of dead plant tissue accumulated during this period, later becoming the fossil fuels.^{[citation needed]}

By feeding on sediments directly to extract the organic component, some detritivores accidentally concentrate toxic pollutants.

Saprophytes [edit]

Saprophyte (-phyte meaning "plant") is a botanical term that is no longer in popular use. ^[4]There are no real saprotrophic organisms that are embryophytes,^{[citation needed]} and fungi and bacteria are no longer placed in the plant kingdom. Plants that were once considered saprophytes, such as non-photosynthetic orchids and monotropes, are now known to be parasites on fungi. These species are now termed myco-heterotrophs.^[5][6][7]

Saprophages [edit]

'Saprophages' are organisms that feed on decaying organic matter.^[8]

See also [edit]

• Decomposer
• Saprotrophic nutrition
• Nepenthes ampullaria
• Consumer-resource systems

References [edit]

v t e Feeding behaviours Carnivores adult Hematophagy Insectivore Lepidophagy Man-eater Molluscivore Mucophagy Myrmecophagy Ophiophagy Piscivore Avivore Spongivore Vermivore Herpetivore reproductive Oophagy Paedophagy Placentophagy Breastfeeding Weaning cannibalistic Cannibalism Human cannibalism Self-cannibalism Sexual cannibalism Herbivores Folivore Frugivore Graminivore Granivore Nectarivore Palynivore Xylophagy Osteophagy Cellular Phagocytosis Myzocytosis Others Bacterivore Fungivore Coprophagia Detritivore Geophagy Omnivore Planktivore Xenophagy Methods Apex predator Bait balls Bottom feeding Browsing Feeding frenzy Filter feeding Grazing Hypercarnivore Intraguild predation Kleptoparasitism Scavenging Trophallaxis Predation Antipredator adaptation Carnivorous plant Carnivorous fungus Carnivorous protist Category:Eating behaviors

v t e Modelling ecosystems – trophic components General Abiotic component Abiotic stress Behaviour Biogeochemical cycle Biomass Biotic component Biotic stress Carrying capacity Competition Ecosystem Ecosystem ecology Ecosystem model Keystone species List of feeding behaviours Metabolic theory of ecology Productivity Producers Autotrophs Chemosynthesis Chemotrophs Foundation species Mixotrophs Myco-heterotrophy Mycotroph Organotrophs Photoheterotrophs Photosynthesis Photosynthetic efficiency Phototrophs Primary nutritional groups Primary production Consumers Apex predator Bacterivore Carnivores Chemoorganotroph Foraging Generalist and specialist species Intraguild predation Herbivores Heterotroph Heterotrophic nutrition Insectivore Mesopredator release hypothesis Omnivores Optimal foraging theory Predation Prey switching Decomposers Chemoorganoheterotrophy Decomposition Detritivores Detritus Microorganisms Archaea Bacteriophage Environmental microbiology Lithoautotroph Lithotrophy Microbial cooperation Microbial ecology Microbial food web Microbial intelligence Microbial loop Microbial mat Microbial metabolism Phage ecology Food webs Biomagnification Ecological efficiency Ecological pyramid Energy flow Food chain Trophic level Example webs Cold seeps Hydrothermal vents Intertidal Kelp forests Lakes North Pacific Subtropical Gyre Rivers San Francisco Estuary Soil Tide pool Processes Ascendency Bioaccumulation Cascade effect Climax community Competitive exclusion principle Consumer-resource systems Copiotrophs Dominance Ecological network Ecological succession Energy quality Energy Systems Language f-ratio Feed conversion ratio Feeding frenzy Mesotrophic soil Nutrient cycle Oligotroph Paradox of the plankton Trophic cascade Trophic mutualism Trophic state index Defense/counter Animal coloration Antipredator adaptations Camouflage Deimatic behaviour Herbivore adaptations to plant defense Mimicry Plant defense against herbivory Predator avoidance in schooling fish

v t e Modelling ecosystems – other components Population ecology Abundance Allee effect Depensation Ecological yield Effective population size Intraspecific competition Logistic function Malthusian growth model Maximum sustainable yield Overpopulation in wild animals Overexploitation Population cycle Population dynamics Population modeling Population size Predator–prey equations Recruitment Resilience Small population size Stability Species Biodiversity Density-dependent inhibition Ecological effects of biodiversity Ecological extinction Endemic species Flagship species Gradient analysis Indicator species Introduced species Invasive species Latitudinal gradients in species diversity Minimum viable population Neutral theory Occupancy-abundance relationship Population viability analysis Priority effect Rapoport's rule Relative abundance distribution Relative species abundance Species diversity Species homogeneity Species richness Species distribution Species-area curve Umbrella species Species interaction Antibiosis Biological interaction Commensalism Community ecology Ecological facilitation Interspecific competition Mutualism Storage effect Spatial ecology Biogeography Cross-boundary subsidy Ecocline Ecotone Ecotype Disturbance Edge effects Foster's rule Habitat fragmentation Ideal free distribution Intermediate Disturbance Hypothesis Island biogeography Landscape ecology Landscape epidemiology Landscape limnology Metapopulation Patch dynamics r/K selection theory Source–sink dynamics Niche Ecological niche Ecological trap Ecosystem engineer Environmental niche modelling Guild Habitat Marine habitats Limiting similarity Niche apportionment models Niche construction Niche differentiation Other networks Assembly rules Bateman's principle Bioluminescence Ecological collapse Ecological debt Ecological deficit Ecological energetics Ecological indicator Ecological threshold Ecosystem diversity Emergence Extinction debt Kleiber's law Liebig's law of the minimum Marginal value theorem Thorson's rule Xerosere Other Allometry Alternative stable state Balance of nature Biological data visualization Constructal theory Ecocline Ecological economics Ecological footprint Ecological forecasting Ecological humanities Ecological stoichiometry Ecopath Ecosystem based fisheries Endolith Evolutionary ecology Functional ecology Industrial ecology Macroecology Microecosystem Natural environment Regime shift Systems ecology Theoretical ecology List of ecology topics